This religious border was designed by German artist Hans Holbein in 1524. It has religious connotations with a bearded man on each side with a halo, and an angel on the top left corner reading. All the animals around the border have halos as well.
Crane, Walter The Decorative Illustration of Books Old and New (New York, NY: G. Bells and Sons, Ltd, 1905)
Courtesy the private collection of Roy Winkelman